Physiological Systems in Insects discusses the roles of molecular biology, neuroendocrinology, biochemistry, and genetics in our understanding of insects. All chapters in the new edition are updated, with major revisions to those covering swiftly evolving areas like endocrine, developmental, behavioral, and nervous systems. The new edition includes the latest details from the literature on hormone receptors, behavioral genetics, insect genomics, neural integration, and much more. Organized according to insect physiological functions, this book is fully updated with the latest and foundational research that has influenced understanding of the patterns and processes of insects and is a valuable addition to the collection of any researcher or student working with insects.

There are about 10 quintillion insects in the world divided into more than one million known species, and some scientists believe there may be more than 30 million species. As the largest living group on earth, insects can provide us with insight into adaptation, evolution, and survival. The internationally respected third edition of Marc Klowden's standard reference for entomologists and researchers and textbook for insect physiology courses provides the most comprehensive analysis of the systems that make insects important contributors to our environment.

Table of Contents


Preface to the Third Edition


Chapter 1. Signaling Systems

Paracrine Signaling

Endocrine Signaling

Types of Hormone Release Sites in Insects

Early Experiments that Set the Stage for Our Current Understanding

Types of Hormones in Insects

Prothoracicotropic Hormone


The Juvenile Hormones

Other Neuropeptides Found in Insects

Terms for Understanding Signaling Systems


Chapter 2. Integumentary Systems

Insect Growth and Development

Strategies for Growth

Instars, Stadia, and Hidden Phases

Structure of the Integument

Modified Features of the Integument

Chemistry of the Cuticle

The Molting Process

Imaginal Disc Development

Metamorphosis, Cell Reorganization, and Cell Death

Terms for Understanding Integumentary Systems


Chapter 3. Developmental Systems

Morphogens and Developmental Patterns

Insect Eggs

Embryonic Development

Terms for Understanding Developmental Systems


Chapter 4. Reproductive Systems

Female Reproductive Systems

Types of Ovarioles


Endocrinology of Female Reproduction

Ovulation, Fertilization and Oviposition

Male Reproductive Systems


Unconventional Methods of Insect Reproduction

Mating Systems

Terms for Understanding Reproductive Systems


Chapter 5. Behavioral Systems

Ways of Looking at Behavior

Physiology of Learning and Memory

Physiology of Behaviors Accompanying Metamorphosis

Behavioral Modification by Parasites

Terms for Understanding Behavioral Systems


Chapter 6. Metabolic Systems

The Insect Alimentary Tract


No. of pages:
© 2013
Academic Press
Print ISBN:
Electronic ISBN:


"This book has grown considerably since its inception in 2002, when it was just over 400 pages in length, due to the addition of new material to all chapters and major revisions to those dealing with rapidly evolving areas such as endocrinology, development, behavior and neurobiology…Klowden has written a very good book, incorporating new findings with long-established principles in the field of insect physiology."--Bulletin of the Entomological Society of Canada, December 2013
"This textbook from Klowden covers insect physiology with both fidelity to the original sites of discovery — often Drosophila — and notes on generalizability to other phyla…Figures are employed to elucidate the anatomical locations and chemistry of the processes discussed."--Reference & Research Book News, October 2013